mature achenes subreniform, ca. 1 mm in diam. , rugose.

capsule rugose, without wings, echinate or lobed, opening by 4 or 5 lobes at maturity.

蒴果具皱纹，没有翅，具刺或浅裂的，以成熟时4或5裂片打开。

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seeds subglobose, ca. 1 mm in diam. , conspicuously ridged, rugose.

近球形的种子，直径约1毫米，显著使成脊状，具皱纹。

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fruit rugose, without prickles and wings, not globose, opening by 4 or 5 lobes at maturity.

果具皱纹，没有皮刺和翅，不球状，成熟时以4或5裂片打开。

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lateral veins 8-18 pairs, appearing rugose due to depressed veins.

侧脉8-18对，由于下凹的脉似乎具皱纹。

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leaf blade veins adaxially deeply impressed, surface rugose; bracteoles and sepals 12-17, inside velutinous.

正面深深凹陷的叶片脉，表面具皱纹；小苞片和萼片12-17，里面短绒毛。

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seeds convex on 1 face, transversely rugose, with fleshy funicle.

种子凸的在1边，横向具皱纹，具肉质的珠柄。

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seeds brown to black-brown, densely rugose.

种子棕色到黑色棕色，浓密。

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seeds deltoid-obconical, ca. 3 mm, densely transversely rugose.

种子正三角形倒圆锥形，约3毫米，浓密横向具皱纹。

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seed coat transversely rugose on dorsal side.

在背侧的边上的横向的种皮具皱纹。

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leaf blade adaxially rugose, abaxially with brown to yellowish brown stellate hairs, margin entire or with scattered teeth.

叶片正面具皱纹，背面具使到淡黄色成褐色棕色毛，边缘全缘或具星散的齿。

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leaf blade oblong, ovate-oblong, or ovate, apex acute, adaxially distinctly rugose.

叶片长圆形，卵状长圆形，或卵形，先端锐尖，正面明显具皱纹。

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seeds curved, rugose, endosperm absent.

种子弄弯，具皱纹，无胚乳。

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seeds brown, ovoid, ca. 3 mm, angled, reticulate-striate, sometimes rugose, apex acuminate.

种子棕色，卵球形，长约3毫米，角，具网状条纹，有时具皱纹，先端渐尖。

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petals white, rarely yellow; achene transversely rugose on each side.

花瓣白色，很少黄；瘦果横向具皱纹在每边。

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young branchlets, leaves, and valves puce when dry; leaf blade papery to thinly leathery; valves rugose and curly when fruit ripe.

干燥时的幼枝，叶和裂爿紫褐色；叶片纸质的到薄革质；裂爿具皱纹和卷曲的果成熟时。

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utricles exceeding perianth, compressed-ovoid, ca. 3 mm, indehiscent, slightly rugose to nearly smooth.

胞果超过花被，压扁卵球形，约3毫米，不裂，稍具皱纹到近平滑的。

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utricles green, longer than perianth, globose, slightly compressed, ca. 2 mm in diam. , very rugose, indehiscent.

绿色的胞果，长于花被，球状，稍压扁，直径约2毫米，具皱纹，不裂。

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style persistent on young fruit, up to 1. 5 mm, valves slightly rugose.

花柱在幼果上宿存，可达1.5毫米，裂爿稍具皱纹。

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young branchlets, leaves, and valves yellowish green when dry; leaf blade thickly leathery; valves not rugose and curly when fruit ripe.

干燥时的幼枝，叶和裂爿黄绿色；叶片厚革质；裂爿不具皱纹和卷曲的果成熟时。

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leaf blade leathery or sometimes membranous, margin entire, undulate-dentate, or rugose.

有时的叶片革质或膜质，边缘全缘，波状具牙齿，或具皱纹。

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